A few year have passed since the Deity of Sin's defeat. Despite a few small conflicts here and there, Gamindustri has remained peaceful, and all the CPUs have been living fun, carefree lives. Then one day, Neptune – the CPU of Planeptune – is swept away to another dimension by sheer happenstance. The new dimension she arrives in looks familiar, but this Gamindustri has a nostalgic 80s feel to it… She also learns upon her arrival that a mysterious group called "The Seven Sages" have cast an evil shadow over the lands of this Gamindustri. Neptune's new journey to return home, as well as to protect this brand-new dimension, is just getting started….